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I've always been a cyclist first and a runner second, by necessity because of my job(s). When I stopped racing bicycles I ran marathons and ultras but I never "enjoyed" it; they were just ways to test myself. I don't have any injuries or other conditions that preclude me from running now but I wouldn't run if you pulled a gun on me.
On topic, IMO the big difference between running fitness and cycling fitness are the surges, especially repeated surges. There's a huge range in which cycling occurs, from very low to very high; running seems to have a much narrower range. A good runner on a bike can get gassed after a few times on the front of a paceline, especially if they don't know where to be or how to make their body one with the bike. Many times, they look like a lumberjack chopping wood with their legs.
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