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I dunno what happened during the ride but after about 5 or so miles my knee hurt like a mot*(&!^!!!.
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I suspect it's more than just "fit", it could be a combination of things.
I have gone through this process myself. Replicating the fit on your bikes will not solve all the problems.
You need to be more in tune with your body, your bike, other external factors - and how they all work together to create a certain outcome. And that will come with time, if you are willing to experiment, try different things, analyze, learn as you go.
Fit is only one piece of the puzzle.
I know you are not a newbie. Just sharing my perspective.